In some cases, the larger cells may be further distinguished based on their scattering characteristics (coccolithophorids) or the presence of both PE and chlorophyll (cryptophytes) (Olson et al., 1989; Collier & Campell, 1999). Many authors have reported the distributions and dynamics of each photosynthetic group in the water column in different marine environments (Li, 1995; Campbell & Vaulot, 1993; Vaulot & Marie, 1999). As both cyanobacteria and picoeukaryotes are widely distributed in the marine environment, they play a fundamental role in aquatic ecosystems because of their high productivity
