LIFE IN EARTH’S ATMOSPHERE presents a formidable challenge to land plants. On the one hand, the atmosphere is the source of carbon dioxide, which is needed for photosynthesis. Plants therefore need ready access to the atmosphere. On the other hand, the atmosphere is relatively dry and can dehydrate the plant. To meet the contradictory demands of maximizing carbon dioxide uptake while limiting water loss, plants have evolved adaptations to control water loss from leaves, and to replace the water lost to the atmosphere. In this chapter we will examine the mechanisms and driving forces operating on water transport within the.