Wild soybean (Glycine soja) is a highly adaptive plant species which can grow well in saline-alkaline soils. In soybean genome, there exist about 140 HD-Zip (Homeodomain-leucine Zipper) genes. HD-Zip transcription factor family is one of the largest plant specific superfamilies and plays important roles in response to abiotic stresses. | A novel Glycine soja homeodomain-leucine zipper HD-Zip I gene Gshdz4 positively regulates bicarbonate tolerance and responds to osmotic stress in Arabidopsis